Pomatoschistus quagga
   (Heckel 1840)

Common names: English: Quagga goby
German: Quagga-Grundel

Common synonyms: none.

Length: 3 - 4 cm.

Yellowish with dark spots on head and back. Laterally 4 (males) or 3 (females) dark bands across. Triangular dark spot on base of caudal fin. Mature females with yellow belly (ripe eggs shining through). Triangular dark bars beween eys and mouth. No black spot in D1.

Large eyes. Caudal fin slightly emarginate (otherwise only in P. knerii). Praedorsal area and breast naked.
35-40 scales in lateral series.
Fins: D1 VI (VI-VII); D2 I+9; A I+9 (8-9); P 15.


Spawning season April to July.

2 - 120 m. Sandy bottoms and sea grass. Free swimming 10 - 50 cm above the bottom. In danger they flee to the ground. Common to very common in some regions.

Pomatoschistus quagga
Photograph: Robert A. Patzner ©
Northern Adriatic, 4 m depth, 30 cm above the bottom, June 1998


Species list
General info


Mediterranean: West-Mediterranean, Adriatic Sea, Aegean Sea; Atlantic.





Literature on Pomatoschistus quagga

De Buen E., 1923: Gobius de la Peninsula Iberica y Baleares. Grupos Lesueurii, Colonianus, Affinis y Minutus. Mem. Inst. eso. Oceonaogr. 3: 121-266.

Fricke, R., M. Bilecenoglu & H. Musa Sari. 2007. Annotated checklist of fish and lamprey species (Gnathostomata and Petromyzontomorphi) of Turkey, including a Red List of threatened and declining species. Stuttgarter Beiträge zur Naturkunde Serie A (Biologie) 706: 1-169.

Heckel J.J., 1840: Ichthyologische Beiträge zu den Familien der Cottiden, Scorpaenoiden, Gobioiden und Cyprinoiden. Ann. naturhist Mus. Wien 2: 145-164.

Kovacic M., 2005: An annotated checklist of the family Gobiidae in the Adriatic Sea. Annales Ser. hist. nat. 15: 1-24.

Kovacic M., 2020: Checklist of gobies (Teleostei: Gobiidae) of the Mediterranean Sea and a key for species identification. Zootaxa 4877: 075-101.

Kovacic M. & Patzner R.A., 2011: North-Eastern Atlantic and Mediterranean Gobies. In: The Biology of Gobies. (Eds: Patzner R.A. et al.) Science Publishers: 177-206.

Kramer A., Kovacic M. & Patzner R.A., 2012: Dentition of eight species of Mediterranean Sea Gobiidae: do dentition characters of gobies reflect phylogenetic relationships? Journal of Fish Biology 80: 29-48.

Lombarte A., Miletic M. & Kovacic M. et al., 2014: Identifying Mediterranean Sea gobies using otoliths. 5th Int. symp. fish otolith res. appl., Peguera, Mallorca: Poster.

Louisy P., 2002: Meeresfische, Westeuropa, Mittelmeer. Ulmer, Stuttgart.

Miller P.J., 1986: Gobiidae. In: Fishes of the North-eastern Atlantic and the Mediterranean. Volume III. (Eds.: Whitehead P.J.P. et al.) pp. 1019-1085. UNESCO, Paris.

Patzner R.A. & H. Moosleitner, 1999: Die Grundeln des Mittelmeeres. Teil 5. Kleine Grundeln des Weichbodens 2. Das Aquarium. 370: 54-60.

Zander C.D., 1982: Zur Morphologie und Biologie einiger seltener Grundeln des Mittelmeeres (Pisces, Gobioidei, Gobiidae). Senck. marit. 14: 1-8.

Zander C.D., 1983: Feeding ecology of littoral gobiid and blennioid fish of the Banyuls area (Mediterranean Sea). 1. Main food and trophic dimension of niche and ecotope. Vie Milieu 32: 1-10.







Photographs of Pomatoschistus quagga

Pomatoschistus quagga
Adriatic Sea, Raab, May 2019
Photograph: Roberto Pillon ©
Pomatoschistus quagga
Adriatic Sea, Raab, May 2019
Photograph: Roberto Pillon ©
Pomatoschistus quagga
Adriatic Sea, Raab, May 2019
Photograph: Roberto Pillon ©
Pomatoschistus quagga
Adriatic Sea, Raab, May 2019
Photograph: Roberto Pillon ©




Drawings of Pomatoschistus quagga


Drawings from Miller (1986)

Copyright © 2019 - Robert A. Patzner, Department of Biosciences, Univ. Salzburg, Austria.
State: 2019-06-13
E-mail: robert.patzner@sbg.ac.at